As the early June sun begins to set over a calm river in Central Hungary, masses of ghostly shapes emerge from their larval cases to take to the air for the first time. They are mayflies and in a spectacular display, thousands of them demonstrate how the very first wings were used. From the stunning aerobatics of hoverflies in an English garden to the mass migration of purple crow butterflies in the valleys of Taiwan, this episode tells the tale of the first animals ever to take to the air. Unique footage reveals the lightning fast reactions of bluebottles and hoverflies, filmed with one of the world's fastest cameras, and Sir David Attenborough handles the world's largest (and perhaps most ferocious) insect - the Titan beetle.

Professor Brian Cox travels from the fossils of the Burgess Shale to the sands of the oldest desert in the world to show how light holds the key to our understanding of the whole universe, including our own deepest origins. To understand how light holds the key to the story of the universe; you first have to understand its peculiar properties. Brian considers how the properties of light that lend colour to desert sands and the spectrum of a rainbow can lead to profound insights into the history and evolution of our universe. Finally, with some of the world's most fascinating fossils in hand, Brian considers how but for an apparently obscure moment in the early evolutionary history of life, all the secrets of light may have remained hidden. Because although the universe is bathed in light that carries extraordinary amounts of information about where we come from, it would have remained invisible without a crucial evolutionary development that allowed us to see. Only because of that development can we now observe, capture and contemplate the incredible wonders of the universe that we inhabit.

Comedian Ben Miller returns to his roots as a physicist to try to answer a deceptively simple question: what is one degree of temperature? His quest takes him to the frontiers of current science as he meets researchers working on the hottest and coldest temperatures in the universe, and to a lab where he experiences some of the strangest effects of quantum physics - a place where super-cooled liquids simply pass through solid glass. Plus, Ben installs his very own Met office weather station at home. Ben's investigations in this personal and passionate film highlight the importance of measurement and accuracy in the 21st century.

In this programme, Mary descends into the city streets to discover the dirt, crime, sex and slum conditions in the world's first high-rise city. This Rome is not the marble Rome we know, but a vast, messy metropolis with little urban planning, where most Romans lived in high-rise apartment blocks with little space, light, or even sanitation. Forced outdoors into the city streets, she reveals where they went to hang out, get drunk, have sex and get clean. She looks at the Forum as a place of gamblers, dentists and thieves, and she explores the lustiness of Roman bar life and jokes. Finally, exploring law and order from the bottom up, Mary examines how this city really worked. She meets Ancarenus Nothus, an apartment dweller who lived in fear of the rent collector; Notorious Primus, who wrote about his three great pleasures in life - baths, wine and sex; and Unlucky Doris, a seven-year-old girl killed in one of Rome's many fires.

With optional Hebrew subtitles. This episode explores how the spread of the Enlightenment brought ghetto walls around Europe crashing down and allowed Jews to join the wider fabric of modern life in Europe in unprecedented ways. This Jewish renaissance saw Giacomo Meyerbeer and Felix Mendelssohn to establish the enduring tradition for Jewish musical prodigies. However the integration of Jewish talent into the mainstream of European culture and commerce eventually stirred up ancient prejudice, expressed in the new fashion of Romantic nationalism and the pseudo-science of anti-semitism.

Life of the Buddha is a major new landmark documentary following Buddha on his journey from the lap of luxury to the verge of starvation and final enlightenment. Shot on location in Nepal and India, Life of the Buddha uses dramatic computer-generated images and recent archaeological discoveries to piece together this remarkable story.