Grasslands cover one quarter of all land and support the vast gatherings of wildlife, but to survive here animals must endure the most hostile seasonal changes on the planet. From Asia's bizarre-looking Saiga antelope to the giant anteaters of Brazil, grassland animals have adapted in extraordinary ways to cope with these extremes. In the flooded Okavango, lions take on formidable buffalo in epic battles, on the savannah bee-eaters take advantage of elephants to help catch insects and, on the freezing northern tundra, caribou embark on great migrations shadowed by hungry Arctic wolves.

The Southern Ocean, which circles the globe without being blocked by land, is home to the Antarctic Circumpolar Current (ACC), the longest of the world's ocean currents. Also known as the "channel", it connects the Indian, Atlantic and Pacific Ocean basins and exerts a powerful influence over the Earth's climate. The ACC carries 150 times more water around Antarctica than the flow of all the world's rivers combined.

In the fourth episode, leading landscape photographer Peter Eastway follows in the footsteps of Frank Hurley – the pioneering Australian photographer on Shackleton’s expedition to the south a century ago – to capture the wilderness and wildlife of Antarctica and South Georgia.

In this final episode we complete our journey, travelling back from the March equinox to the end of June. Kate Humble is in the Arctic at a place where spring arrives with a bang, whilst Helen Czerski chases a tornado to show how the earth's angle of tilt creates the most extreme weather on earth.

The polar regions of our planet may seem beyond the reach of most of us, but they are not beyond our influence. We, unintentionally, are changing these frozen worlds, and these changes will not just affect the poles but the whole planet. On the unforgiving frontier of climate change, polar bears, walruses, seals and penguins find their icy Edens in peril.

Every year, around the world, seasonal changes transform entire landscapes and draw in millions of creatures as these great events unfold. This fantastic series combines the epic scale of Planet Earth and the intimate, emotional stories of charismatic animals as they struggle to survive. Using state of the art technology, these programmes capture the Earth's most dramatic and epic wildlife spectacles and the intimate stories of the animals caught up in them.
The Great Melt: The summer melt of Arctic ice, opening up nearly three million square miles of ocean and land, provides opportunities for millions of animals, including beluga whales, families of Arctic foxes, vast colonies of seabirds, and the fabled Arctic unicorn, the narwhal. For polar bears, however, it is the toughest time of year. Why? How will they survive? A mother polar bear and her cub make their first journey together onto the sea ice. They are looking for ringed seals, their favourite prey. It is a serious business but the cub just wants to play. The melting ice makes it harder for them to hunt and threatens their survival. In a unique aerial sequence, the migration of narwhal with their distinctive unicorn-like tusks is filmed for the first time. The whales' journey is risky as they travel along giant cracks in the ice. If the ice were to close above them, they would drown. Hundreds of beluga whales gather in the river shallows. They rub themselves on smooth pebbles in one of the most bizarre summer spectacles. Guillemot chicks take their first flights from precipitous sea cliff nests to the sea 300 metres below. They attempt to glide to safety but many miss their target. Their loss is a bonus for the hungry Arctic fox family waiting below. As the melt comes to an end the bears gather, waiting for the sea to freeze again. Two 400kg males square up to each other to spar.