In the far north of our planet lies the largest land habitat on earth, home to snow-covered forests and the icy open tundra. These are lands of extremes that push animals to their limits: in winter they are so cold that much of the ground has remained frozen since the last ice age. To stand any chance of survival, animals must adapt in extreme ways: here a super pack of wolves, 25 strong, has come together to take on the only large prey available to them in winter, American bison.
On the featureless tundra, an Arctic fox must strike a living alone. She is a wanderer and will roam many hundreds of miles searching for tiny lemmings, hidden deep underground. The only way to reach them is with a head dive. In the remote far east of Russia, a rare Amur leopard prowls the seemingly empty, snow-covered forest. With little prey available, it must use its ingenuity to find a meal. It follows crows in the hope of finding carrion, but it must not stay long, for it shares the forest with a far larger but equally hungry big cat, the Siberian tiger.
As spring arrives, the forests begin to thaw and life returns. Beneath the ground, a nest of tiny painted turtle hatchlings now emerge, having remained frozen in a state of suspended animation throughout winter. To the north, it is a further month before the sun’s warmth baths the frozen ground of the tundra. Tucked away underground lies a tiny snow queen – a Lapland bumble bee. She is the sole survivor of her colony - the rest perished in the winter freeze - but her larger size, her furry body and antifreeze in her blood have allowed her to survive. Now she is in a hurry. She must feed herself and raise a brood in the brief window of summer while the flowers are in bloom.
Snowy owls also use the open tundra to breed: one pair have raised a nest full of fluffy chicks. With 24-hour daylight in which to hunt, the dedicated parents bring back meal after meal for their ever-growing brood. But one day, they return to find the nest empty.
Today, the biggest challenge in the tundra is climate change. Warming summers are melting the permafrost deep within the soil, causing the ground to thaw and, in places, the land to collapse. These changes are impacting the animals too. Caribou arrive in herds of 200,000 individuals to raise their calves in the rich pastures, but warming means mosquitos emerge sooner and bother the calves before they have had a chance to gain strength. The parents drive their young to cooler, mosquito-free land, but to get there they must cross rivers running with increased meltwater and escape hungry grizzly bears. They, like much of the tundra's wildlife, are adapted to live in the extremes - but the challenge of today’s warming climate could be one extreme too many.

At the top of our planet lies a magical realm, the Arctic Ocean. After four months of winter darkness, the sun returns to reveal a frozen ocean covered in ice. Mother polar bears emerge from their hillside dens and lead their cubs down to the sea ice to hunt, while a young male and female bear forge a surprising friendship out on the ice.
For others, the frozen sea is a trap. A pod of beluga whales has been confined to an ice hole for five months, slowly starving to death as the food around them runs out. Their salvation lies in the strengthening sun that comes with spring, melting the sea ice, allowing their escape.
Off the east coast of Greenland, the floating pack ice in spring is a nursery ground for harp seals. Mothers and pups have just a few weeks together for the pup to learn to swim before she leaves him to fend for himself. But in today’s warming climate, storms can tip helpless youngsters into the sea before they are strong enough to fend for themselves.
Summer is a time of plenty in the Arctic Ocean as plankton blooms feed millions of tiny mouths, such as bizarre skeleton shrimps, as well as the biggest: bowhead whales. These ancient and long-lived whales arrive en masse every year at secret locations known as whale spas. But today, with the loss of summer sea ice, their peace is shattered by orcas from the south. These daring predators are bold enough to take on the much larger bowheads, targeting their vulnerable calves.
The 24-hour daylight of the Arctic summer attracts visitors from afar, including huge flocks of seabirds like crested auklets. A male must use both his song and a secret tangerine perfume if he is to attract a mate. For the resident walrus, the summer heat can be unbearable. After hauling himself to the beach to moult, an old male uses an ingenious technique to get himself back to the cool of the water - a roly-poly!
Summers in the Arctic today bring record-breaking heat. With climate change, it is warming faster than anywhere else on Earth. It is predicted that the Arctic Ocean could become ice-free each summer by 2035, raising new challenges for polar bears. Without sea ice, more and more bears are becoming stranded on remote Arctic islands. It's a dangerous place to be for a mother bear with cubs, surrounded by larger, predatory males.

Sir David Attenborough journeys to both Polar Regions to investigate what rising temperatures will mean for the people and wildlife that live there and for the rest of the planet. David starts out at the North Pole, standing on sea ice several metres thick, but which scientists predict could be Open Ocean within the next few decades. The Arctic has been warming at twice the global average, so David heads out with a Norwegian team to see what this means for polar bears. He comes face-to-face with a tranquilised female, and discovers that mothers and cubs are going hungry as the sea ice on which they hunt disappears. In Canada, Inuit hunters have seen with their own eyes what scientists have seen from space; the Arctic Ocean has lost 30% of its summer ice cover over the last 30 years. For some, the melting sea ice will allow access to trillions of dollars worth of oil, gas and minerals. For the rest of us, it means the planet will get warmer, as sea ice is important to reflect back the sun's energy. Next David travels to see what's happening to the ice on land: in Greenland, we follow intrepid ice scientists as they study giant waterfalls of meltwater, which are accelerating iceberg calving events, and ultimately leading to a rise in global sea level. Temperatures have also risen in the Antarctic - David returns to glaciers photographed by the Shackleton expedition and reveals a dramatic retreat over the past century. It's not just the ice that is changing - ice-loving adelie penguins are disappearing, and more temperate gentoo penguins are moving in. Finally, we see the first ever images of the largest recent natural event on our planet - the break up of the Wilkins Ice Shelf, an ice sheet the size of Jamaica, which shattered into hundreds of icebergs in 2009.

Spring arrives in the polar regions, and the sun appears after an absence of five months; warmth and life return to these magical ice worlds - the greatest seasonal transformation on our planet is underway. Male Adelie penguins arrive in Antarctica to build their nests - it takes a good property to attract the best mates and the males will stop at nothing to better their rivals! But these early birds face the fiercest storms on the planet. In the Arctic, a polar bear mother is hunting with her cubs. Inland, the frozen rivers start to break up and billions of tons of ice are swept downstream in the greatest of polar spectacles. This melt-water fertilizes the Arctic Ocean, feeding vast shoals of Arctic cod and narwhal. The influx of freshwater accelerates the breakup of the sea-ice - an area of ice the size of Australia will soon vanish from the Arctic. On land, a woolly bear caterpillar emerges from the snow having spent the winter frozen solid. Caterpillars normally become moths within months of hatching, but life is so harsh here that the woolly bear takes 14 years to reach adulthood. Once mature it has only days to find a mate before it dies! Alongside the caterpillars white Arctic wolves race to raise their adorable cubs before the cold returns.

It is high summer in the Polar Regions, and the sun never sets. Vast hordes of summer visitors cram a lifetime of drama into one long, magical day; they must feed, fight and rear their young in this brief window of plenty. Summer is a tough time for the polar bear family, as their ice world melts away and the cubs take their first swimming lesson. Some bears save energy by dozing on icy sun beds, while others go egg-collecting in an Arctic tern colony, braving bombardment by sharp beaks. There are even bigger battles on the tundra; a herd of musk oxen gallop to the rescue as a calf is caught in a life and death struggle with a pair of Arctic wolves. But summer also brings surprises, as a huge colony of 400,000 king penguins cope with an unlikely problem - heat. The adults go surfing, while the woolly-coated chicks take a cooling mud bath. Nearby, a bull fur seal is prepared to fight to the death with a rival. Fur flies as the little pups struggle desperately to keep out of the way of the duelling giants. Further south, a minke whale is hunted amongst the ice floes by a family of killer whales. The dramatic chase lasts over 2 hours and has never been filmed before. The killers harry the minke whale, taking it in turns to wear it down. Eventually it succumbs to the relentless battering. Finally, comical adelie penguins waddle back to their half a million strong colony like clockwork toys. The fluffy chicks need constant feeding and protection as piratical skuas patrol the skies. When an unguarded chick is snatched, a dramatic "dogfight" ensues.

Our journey begins with David Attenborough at the North Pole, as the sun returns after six months of darkness. We follow a pair of courting polar bears, which reveal a surprisingly tender side. Next stop is the giant Greenland ice cap, where waterfalls plunge into the heart of the ice and a colossal iceberg carves into the sea. Humpback whales join the largest gathering of seabirds on earth to feast in rich Alaskan waters. Further south, the tree line marks the start of the Taiga forest, containing one third of all trees on earth. Here, 25 of the world's largest wolves take on formidable bison prey. At the other end of our planet, the Antarctic begins in the Southern Ocean where surfing penguins struggle to escape a hungry sea-lion and teams of orcas create giant waves to wash seals from ice floes -a filming first. Diving below the ice, we discover prehistoric giants, including terrifying sea spiders and woodlice the size of dinner plates. Above ground, crystal caverns ring the summit of Erebus, the most southerly volcano on earth. From here we retrace the routes of early explorers across the formidable Antarctic ice-cap - the largest expanse of ice on our planet. Finally, we rejoin David at the South Pole, exactly one hundred years after Amundsen then Scott were the first humans to stand there