East Africa is a land which is constantly changing. To survive here, creatures must be able to deal with unpredictable twists and turns - wet turning to dry, feast to famine, cold to hot - no matter how hostile it becomes. From dense forests to snow capped peaks, steamy swamps and endless savannah, this unique and varied land is also a haven for life, supporting large animals in numbers found nowhere else on Earth. But away from the familiar, forever-travelling herds, there are a huge cast of other characters - lizards that steal flies from the faces of lions, vast dinosaur-like birds who stalk catfish through huge wetlands, and an eagle who risks everything on the arrival of ten million bats from a far off rainforest.

David Attenborough discovers the plants that have evolved to shed their dependency on water enabling them to survive in the driest environments. The story begins at midnight in midsummer as David steps into the Princess of Wales Conservatory to witness the extraordinary nocturnal blooming of a cactus. The queen of the night, with its giant flowers, is the centre piece of a stunning symphony of cacti blooms that burst open in the desert (and at Kew) at night. In a mesmerizing 3D slow motion sequence, we discover the extraordinary connections between cacti and their natural pollinators: bats. As the sun rises, David meets other amazing plants. Species like the century plant, the Agave franzosini, which grows steadily for over 50 years, only to then flower itself to death with one mighty telegraph pole sized bloom which literally bursts out of the roof of Kew’s green house.

The first episode looks at how birds first took to the skies in the wake of the insects. It begins in Mexico, where Sir Attenborough observes bats being outmanoeuvred by a red-tailed hawk. Pterosaurs were the birds' forerunners, some 150 million years after dragonflies developed the means of flight, but eventually went extinct together with the dinosaurs. Birds had by then already evolved from early forms like archaeopteryx, the first creature to possess feathers. Its ancestry can be traced through reptiles, and some current species, such as the flying lizard, possibly show paths this evolution may have taken." One of the biggest birds to have ever existed was the terror bird, which proliferated after dinosaurs vanished and stood up to 2.5 metres tall. By comparison, the ostrich, while not closely related, is the largest and heaviest living bird. It was probably the evasion of predators that drove most birds into the air, so their flightless cousins evolved because they had few enemies. Accordingly, such species are more likely to be found on islands, and Sir Attenborough visits New Zealand to observe its great variety, most especially the kiwi. Also depicted is the moa, another huge creature that is now gone. The takahē is extremely rare, and high in the mountains of New Zealand, Sir Attenborough discovers one from a population of only 40 pairs. Finally, another example on the brink of extinction is the kakapo, which at one point numbered only 61 individuals. A male is heard calling — an immensely amplified deep note that can be heard at great distances from its nest.

David Attenborough concludes his epic history of the evolution of flight with an exploration of the highly advanced fliers that dominate our skies today – the extraordinarily diverse skills of the birds, and the sonar-guided precision of the bats. He encounters some of our planet’s most remarkable fliers – from peregrine falcons dive-bombing starlings over Rome, to hummingbirds hovering in the cloud forests of Ecuador. In a spectacular finale, he visits Gomantong cave in Borneo to witness the mass exodus of a million bats.